Boon, C.R. & Outen, A.R. 2011. Flora of Bedfordshire. Bedfordshire Natural History Society. The latest in a line of comprehensive Bedfordshire floras from 1798 onwards.

Laflin, T. 1953. Musci. In: Flora of Bedfordshire, by Dony, J.G., pp. 160-190. Luton: Corporation of Luton Museum and Art Gallery.

Outen, A. 1989. The liverworts of Bedfordshire. Bedfordshire naturalist, 43: 7-18.

Taylor, P. 1953. Hepaticae. In: Flora of Bedfordshire by Dony, J.G., pp. 155-159. Luton: Corporation of Luton Museum and Art Gallery.